The British Rail Class 378 Capitalstar[4] is an electric multiple unit passenger train specifically designed for the London Overground network. It is part of Bombardier Transportation's Electrostar family. A total of 57 five-car trains have been built - most of which were originally built as three- or four-car units.

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The Class 378s were ordered in August 2006 in response to passenger dissatisfaction with existing mass-transit trains operated by Silverlink, such as the ageing fleet of Class 313. The procurement of a modern replacement was viewed as yielding better performance than the refurbishment of the existing units. Accordingly, a total of 57 Class 378s were manufactured at Bombardier's Derby site, the first example of which entered revenue service during July 2009. The type's introduction was roughly half a year later than originally planned, largely as a result of difficulties with multiple suppliers.

In design terms the train is broadly similar to the Class 376 trains operated by Southeastern, featuring the same wider metro-style sliding pocket doors for more efficient boarding and alighting. However, it also has significant differences from the Class 376, such as fully longitudinal seating similar to that used on London Underground rolling stock to give more standing and less seating capacity and reduce overcrowding, the interior having been optimised for the high-volume metro-style services being operated by London Overground.
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