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Bony fish, distinguished by the presence of swim bladders and later ossified endoskeletons, emerged as the dominant group of fish after the end-Devonian extinction wiped out the apex predators, the placoderms. Bony fish are further divided into the lobe-finned and ray-finned fish. About 96% of all living fish species today are teleosts, a crown group of ray-finned fish that can protrude their jaws. The tetrapods, a mostly terrestrial clade of vertebrates that have dominated the top trophic levels in both aquatic and terrestrial ecosystems since the Late Paleozoic, evolved from lobe-finned fish during the Carboniferous, developing air-breathing lungs homologous to swim bladders. Despite the cladistic lineage, tetrapods are usually not considered to be fish, making "fish" a paraphyletic group.
Fish have been an important natural resource for humans since prehistoric times, especially as food. Commercial and subsistence fishers harvest fish in wild fisheries or farm them in ponds or in breeding cages in the ocean. Fish are caught for recreation, or raised by fishkeepers as ornaments for private and public exhibition in aquaria and garden ponds. Fish have had a role in human culture through the ages, serving as deities, religious symbols, and as the subjects of art, books and movies.
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