Wild West C.O.W.-Boys of Moo Mesa dealt with a mutation of some kind; an irradiated comet struck the late 19th century Western plains creating a miles high mesa shrouded in clouds. Everything trapped on top of the mesa was "cow-metized" by the light from the "cow-met" and "evolved" into a "bovipomorphic" state.
Inspired by old tales of the Wild West, this new bovine community developed to the point where they emulated that era's way of life, including the requisite ruffians and corrupt sheriffs.

The player characters in the game are The Cowlorado Kid, The Dakota Dude, Marshal Moo Montana, and Buffalo Bull. The game can be configured to allow up to four players to play simultaneously.

The game's controls consist of an eight-way joystick and two buttons for shooting and jumping. Pressing both buttons simultaneously will cause the player character to perform a stampede attack, do a spinning jump, or use any equipped item like dynamite sticks and tomahawk blades. The player can uncover a variety of power-up items by shooting flying chickens, which includes a rapid-fire Machine Gun and a Power-Up for wider bullets. Unlike Sunset Riders, the player has a three-unit life gauge that allows the player to take three hits before losing a life.

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