Once you've learned an ability, you can simply tap it to use it.
Some abilities, such as Ringo's Achille's Shot and Hellfire Brew, are directed at your current target. These abilities won't work if you haven't tapped an enemy to target them first.
Other abilities bring up a ring, indicating that you can select a target or aim the ability by tapping somewhere within the ring. If you've made a mistake, you can cancel these by tapping the ability a second time.
Most abilities require energy before they can be used. Once they're used, they'll go into a cooldown period before they can be used again.
You can increase your hero's effectiveness with abilities by purchasing blue Crystal items. These are found by tapping the "Ability" tab on the left-hand side of the shop.
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